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Subprogram C2: Nitrogen Induced Ecosystem Changes  
Nitrogen Dose - Vegetation Responses  
The goal is to clarify the biochemical mechanisms involved in the interactions between plants in coniferous forests and their natural enemies. This includes to clarify the numerical response of the ground layer plants and to find suitable indicators for assessing nitrogen deposition effects on vegetation. New and original data on dose-response relationships for nitrogen induced ecosystem changes to forest ecosystems will be provided.

Of prime importance is to understand to what extent vegetation effects due to increased nitrogen deposition are scale-dependent. The studies have been able to quantify a drastic increase in the abundance of natural enemies already at low levels of nitrogen supply and preliminary data indicates that these attacks have a measurable impact on species composition. This indicates that the interaction between parasitic fungi, herbivorous insects and vegetation is an important factor regulating species assemblage in coniferous forests.

The studies comprises:

· Additions of different doses of nitrogen in experiments with plot sizes between 1 and 5000 m2.

· Identification of simple indicators that can be used in assessments of the risk for significant effects on species composition in ecosystems subject to nitrogen deposition.

· Development of new concepts of critical load and environmental monitoring (together with other activities of ASTA).

Lars Ericson
Umeå University
lars.ericson@eg.umu.se